Bobbink, R., During, H. J., Schreurs, J., Willems, J., and Zielman, R.:
Effects of Selective Clipping and Mowing Time on Species Diversity in Chalk
Grassland, Folia Geobot. Phytotx., 22, 363–376, 1987.
Bohner, A., Grims, F., Sobotik, M., and Zechner, L.: Die
Trespen-Halbtrockenrasen (Mesobrometum erecti KOch 1926) im mittleren
steirischen Ennstal (Steiermark, Österreich) – Ökologie, Soziologie
und Naturschutz, Tuexenia, 23, 199–226,
2003.
Bohner, A., Starlinger, F., and Koutecky, P.: Vegetation changes in an
abandoned montane grassland, compared to changes in a habitat with
low-intensity sheep grazing – A case study in Styria, Austria, Eco. Mont.,
4, 5–12, https://doi.org/10.1553/eco.mont-4-2s5, 2012.
Bouché, M. B.: Stratégies lombriciennes, in: Soil
Oranism as Components of Ecosystems, Ecol. Bull., 25, 122–132, 1977.
Brohmer, P.: Fauna von Deutschland, Ein Bestimmungsbuch unserer heimischen
Tierwelt, Quelle & Meyer, Heidelberg, 1984.
Brown, G. G.: How do earthworms affect microflora and faunal community
diversity?, Plant Soil, 170, 209–231, 1995.
Christian, E. and Zicsi, A.: Ein synoptischer bestimmungsschlussel der
regenwurmer Osterreichs (Oligochaeta: Lumbricidae), Die Bodenkultur, 50,
121–131, https://doi.org/10.1002/anie.200805838, 1999.
Cluzeau, D., Guernion, M., Chaussod, R., Martin-Laurent, F., Villenave, C.,
Cortet, J., Ruiz-Camacho, N., Pernin, C., Mateille, T., Philippot, L.,
Bellido, A., Rougé, L., Arrouays, D., Bispo, A., and Pérès, G.:
Integration of biodiversity in soil quality monitoring: Baselines for
microbial and soil fauna parameters for different land-use types, Eur. J.
Soil Biol., 49, 63–72, https://doi.org/10.1016/j.ejsobi.2011.11.003, 2012.
Decaëns, T., Dutoit, T., and Alard, D.: Earthworm community
characteristics during afforestation of abandoned chalk grasslands (Upper
Normandy, France), Eur. J. Soil Biol., 33, 1–11, 1997.
Del-Val, E. and Crawley, M. J.: What limits herb biomass in grasslands:
Competition or herbivory?, Oecologia, 142, 202–211,
https://doi.org/10.1007/s00442-004-1719-8, 2005.
Eisenhauer, N., Milcu, A., Sabais, A. C. W., Bessler, H., Weigelt, A.,
Engels, C., and Scheu, S.: Plant community impacts on the structure of
earthworm communities depend on season and change with time, Soil Biol.
Biochem., 41, 2430–2443, https://doi.org/10.1016/j.soilbio.2009.09.001, 2009.
Fischer, M. A., Oswald, K., and Adler, W.: Exkursionsflora für
Österreich, Lichtenstein und Südtirol, 3rd edn., Land
Oberösterreich, Biologiezentrum der Oberösterreichischen
Landesmuseen, Linz, 2008.
Fontana, V., Radtke, A., Walde, J., Tasser, E., Wilhalm, T., Zerbe, S., and
Tappeiner, U.: What plant traits tell us: Consequences of land-use change of
a traditional agro-forest system on biodiversity and ecosystem service
provision, Agric. Ecosyst. Environ., 186, 44–53,
https://doi.org/10.1016/j.agee.2014.01.006, 2014.
Grime, J. P.: Evidence for the Existence of Three Primary Strategies in
Plants and Its Relevance to Ecological and Evolutionary Theory, Am. Nat.,
111, 1169–1194, https://doi.org/10.1086/283244, 1977.
Groffman, P. M., Eagan, P., Sullivan, W. M., and Lemunyon, J. L.: Grass
species and soil type effects on microbial biomass and activity, Plant Soil,
183, 61–67, https://doi.org/10.1007/BF02185565, 1996.
Hättenschwiler, S., Tiunov, A. V., and Scheu, S.: Biodiversity and litter
decomposition in terrestrial ecosystems, Annu. Rev. Ecol. Evol. Syst., 36,
191–218, https://doi.org/10.1146/annurev.ecolsys.36.112904.151932, 2005.
Hejcman, M., Hejcmanová, P., Pavlů, V., and Beneš, J.: Origin and
history of grasslands in Central Europe – A review, Grass Forage Sci.,
68, 345–363, https://doi.org/10.1111/gfs.12066, 2013.
Hooper, D. U., III, F. S. C., Ewel, J. J., Hector, A., Inchausti, P.,
Lavorel, S., Lawton, J. H., Lodge, D. M., Loreau, M., Naeem, S., Schmid, B.,
Setälä, H., Symstad, A. J., Vandermeer, J., and Wardle, D. A.:
Effects of biodiversity on ecosystem functioning: a consensus of current
knowledge, Ecol. Monogr., 75, 3–35, https://doi.org/10.1890/04-0922, 2005.
Hussain, R. I., Walcher, R., Brandl, D., Jernej, I., Arnberger, A., Zaller,
J. G., and Frank, T.: Influence of abandonment on syrphid assemblages in
mountainous meadows, J. Appl. Entomol., 142, 450–456,
https://doi.org/10.1111/jen.12482, 2018.
Ivask, M., Kuu, A., Truu, M., Kutti, S., Meriste, M., and Peda, J.: Earthworm
communities in soils of Estonian wooded meadows, Balt. For., 18, 111–118,
2012.
Jacquemyn, H., Brys, R., and Hermy, M.: Short-term effects of different
management regimes on the response of calcareous grassland vegetation to
increased nitrogen, Biol. Conserv., 111, 137–147,
https://doi.org/10.1016/S0006-3207(02)00256-2, 2003.
Józefowska, A., Zaleski, T., Zarzycki, J., and Frączek, K.: Do mowing
regimes affect plant and soil biological activity in the mountain meadows of
Southern Poland?, J. Mt. Sci., 15, 2409–2421,
https://doi.org/10.1007/s11629-018-4953-y, 2018.
Karrer, J.: Abandonment of nutrient poor montane meadows: impacts on
regulating ecosystem services (decomposition, greenhouse gas efflux) and
plant diversity, University of Natural Resources and Life Sciences, Vienna,
2015.
Kelemen, A., Török, P., Valkó, O., Miglécz, T. and Tóthmérész, B.: Mechanisms shaping plant biomass and species richness: Plant strategies and litter effect in alkali and loess grasslands, J. Veg. Sci., 24, 1195–1203, doi:10.1111/jvs.12027, 2013.
Kelemen, A., Török, P., Valkó, O., Deák, B., Miglécz,
T., Tóth, K., Ölvedi, T., and Tóthmérész, B.: Sustaining
recovered grasslands is not likely without proper management: vegetation
changes after cessation of mowing, Biodivers. Conserv., 23, 741–751,
https://doi.org/10.1007/s10531-014-0631-8, 2014.
Keuskamp, J. A., Dingemans, B. J. J., Lehtinen, T., Sarneel, J. M., and
Hefting, M. M.: Tea Bag Index: a novel approach to collect uniform
decomposition data across ecosystems, Methods Ecol. Evol., 4,
1070–1075, https://doi.org/10.1111/2041-210X.12097, 2013.
Köhler, B., Gigon, A., Edwards, P. J., Krüsi, B., Langenauer, R.,
Lüscher, A., and Ryser, P.: Changes in the species composition and
conservation value of limestone grasslands in Northern Switzerland after 22
years of contrasting managements, Perspect. Plant Ecol. Evol. Syst., 7,
51–67, https://doi.org/10.1016/j.ppees.2004.11.003, 2005.
Lavelle, P.: Biology and ecology of earthworms, 3rd edn., Chapman & Hall,
UK, 1997.
Lavelle, P., Pashanasi, B., Charpentier, F., Gilot, C., Rossi, J.-P.,
Derouard, L., Andre, J., Ponge, J.-F., and Bernier, N.: Large-Scale Effects
of Earthworms on Soil Organic Matter and Nutrient Dynamics, in: Earthworm
Ecology, edited by: C. A. Edwards, St. Lucie Press, 103–122, 1998.
Lieskovský, J., Bezák, P., Špulerová, J., Lieskovský,
T., Koleda, P., Dobrovodská, M., Bürgi, M., and Gimmi, U.: The
abandonment of traditional agricultural landscape in Slovakia – Analysis of
extent and driving forces, J. Rural Stud., 37, 75–84,
https://doi.org/10.1016/j.jrurstud.2014.12.007, 2015.
Liu, L., King, J. S., Booker, F. L., Giardina, C. P., Lee Allen, H., and Hu,
S.: Enhanced litter input rather than changes in litter chemistry drive soil
carbon and nitrogen cycles under elevated CO2: a microcosm study, Glob.
Chang. Biol., 15, 441–453, https://doi.org/10.1111/j.1365-2486.2008.01747.x, 2009.
Lubbers, I. M., Pulleman, M. M., and Van Groenigen, J. W.: Can earthworms
simultaneously enhance decomposition and stabilization of plant residue
carbon?, Soil Biol. Biochem., 105, 12–24,
https://doi.org/10.1016/j.soilbio.2016.11.008, 2017.
MacDonald, D., Crabtree, J. R., Wiesinger, G., Dax, T., Stamou, N., Fleury,
P., Gutierrez Lazpita, J., and Gibon, A.: Agricultural abandonment in
mountain areas of Europe: Environmental consequences and policy response, J.
Environ. Manage., 59, 47–69, https://doi.org/10.1006/jema.1999.0335, 2000.
Marini, L., Klimek, S., and Battisti, A.: Mitigating the impacts of the
decline of traditional farming on mountain landscapes and biodiversity: a
case study in the European Alps, Environ. Sci. Policy, 14, 258–267,
https://doi.org/10.1016/j.envsci.2010.12.003, 2011.
Maurer, K.: Natural and anthropogenic determinants of biodiversity of
grasslands in the Swiss Alps, Universität Basel, 2005.
Meier, C. L. and Bowman, W. D.: Links between plant litter chemistry,
species diversity, and below-ground ecosystem function, P. Natl. Acad.
Sci. USA, 105, 19780–19785, https://doi.org/10.1073/pnas.0805600105, 2008.
Milcu, A., Partsch, S., Scherber, C., Weisser, W. W., and Scheu, S.:
Earthworms and legumes control litter decomposition in a plant diversity
gradient, Ecology, 89, 1872–1882, https://doi.org/10.1890/07-1377.1, 2008.
Moog, D., Poschlod, P., Kahmen, S., and Schreiber, K. F.: Comparison of
species composition between different grassland management treatments after
25 years, Appl. Veg. Sci., 5, 99–106,
https://doi.org/10.1111/j.1654-109X.2002.tb00539.x, 2002.
Niedrist, G., Tasser, E., Lüth, C., Dalla Via, J., and Tappeiner, U.:
Plant diversity declines with recent land use changes in European Alps,
Plant Ecol., 202, 195–210, https://doi.org/10.1007/s11258-008-9487-x, 2009.
Novara, A., Rühl, J., La Mantia, T., Gristina, L., La Bella, S., and Tuttolomondo, T.: Litter contribution to soil organic carbon in the processes of agriculture abandon, Solid Earth, 6, 425–432, https://doi.org/10.5194/se-6-425-2015, 2015.
Pavlů, L., Pavlů, V., Gaisler, J., Hejcman, M., and Mikulka, J.:
Effect of long-term cutting versus abandonment on the vegetation of a
mountain hay meadow (Polygono-Trisetion) in Central Europe, Flora – Morphol.
Distrib. Funct. Ecol. Plants, 206, 1020–1029,
https://doi.org/10.1016/j.flora.2011.07.008, 2011.
Peco, B., Carmona, C. P., de Pablos, I., and Azcárate, F. M.: Effects of
grazing abandonment on functional and taxonomic diversity of Mediterranean
grasslands, Agric. Ecosyst. Environ., 152, 27–32,
https://doi.org/10.1016/j.agee.2012.02.009, 2012.
Piotrowska, K., Connolly, J., Finn, J., Black, A., and Bolger, T.: Evenness
and plant species identity affect earthworm diversity and community
structure in grassland soils, Soil Biol. Biochem., 57, 713–719,
https://doi.org/10.1016/j.soilbio.2012.06.016, 2013.
Pižl, V.: Succession of earthworm populations in abandoned fields, Soil
Biol. Biochem., 24, 1623–1628, https://doi.org/10.1016/0038-0717(92)90160-Y, 1992.
Ponge, J.-F., Pérès, G., Guernion, M., Ruiz-Camacho, N., Cortet, J.,
Pernin, C., Villenave, C., Chaussod, R., Martin-Laurent, F., Bispo, A., and
Cluzeau, D.: The impact of agricultural practices on soil biota: A regional
study, Soil Biol. Biochem., 67, 271–284, https://doi.org/10.1016/j.soilbio.2013.08.026,
2013.
Pop, V. V: Earthworm-Vegetation-Soil Relationships in the Romanian
Carpathians, Soil Biol. Biochem., 29(, 223–229, 1997.
Prévosto, B., Kuiters, L., Bernhardt-Römermann, M., Dölle, M.,
Schmidt, W., Hoffmann, M., Van Uytvanck, J., Bohner, A., Kreiner, D.,
Stadler, J., Klotz, S., and Brandl, R.: Impacts of Land Abandonment on
Vegetation: Successional Pathways in European Habitats, Folia Geobot.,
46, 303–325, https://doi.org/10.1007/s12224-010-9096-z, 2011.
R Core Team: A language and environment for statistical computing, Version v 3.1., Vienna, Austria, available at:
http://www.R-project.org (last access: 23 August 2019), 2016.
Scherer-Lorenzen, M.: Functional diversity affects decomposition processes
in experimental grasslands, Funct. Ecol., 22, 547–555,
https://doi.org/10.1111/j.1365-2435.2008.01389.x, 2008.
Scheu, S.: Changes in the lumbricid coenosis during secondary succession
from a wheat field to a beechwood on limestone, Soil Biol. Biochem., 24,
1641–1646, https://doi.org/10.1016/0038-0717(92)90163-R, 1992.
Schmitzberger, I., Wrbka, T., Steurer, B., Aschenbrenner, G., Peterseil, J.,
and Zechmeister, H. G.: How farming styles influence biodiversity
maintenance in Austrian agricultural landscapes, Agric. Ecosyst. Environ.,
108, 274–290, https://doi.org/10.1016/j.agee.2005.02.009, 2005.
Seeber, J., Scheu, S., and Meyer, E.: Effects of macro-decomposers on litter
decomposition and soil properties in alpin
e pastureland: A mesocosm
experiment, Appl. Soil Ecol., 34, 168–175,
https://doi.org/10.1016/j.apsoil.2006.02.004, 2006.
Spehn, E. M., Joshi, J., Schmid, B., Alphei, J., and Körner, C.: Plant
diversity effects on soil heterotrophic activity in experimental grassland
ecosystems, Plant Soil, 224, 217–230, https://doi.org/10.1023/A, 2000.
Tasser, E. and Tappeiner, U.: Impact of land use changes on mountain
vegetation, Appl. Veg. Sci., 5, 173–184,
https://doi.org/10.1658/1402-2001(2002)005[0173:IOLUCO]2.0.CO;2, 2002.
Walcher, R., Karrer, J., Sachslehner, L., Bohner, A., Pachinger, B., Brandl,
D., Zaller, J. G., Arnberger, A., and Frank, T.: Diversity of bumblebees,
heteropteran bugs and grasshoppers maintained by both: abandonment and
extensive management of mountain meadows in three regions across the
Austrian and Swiss Alps, Landsc. Ecol., 32, 1937–1951,
https://doi.org/10.1007/s10980-017-0556-1, 2017.
Wehn, S., Taugourdeau, S., Johansen, L., and Hovstad, K. A.: Effects of
abandonment on plant diversity in semi-natural grasslands along soil and
climate gradients, J. Veg. Sci., 28, 838–847, https://doi.org/10.1111/jvs.12543,
2017.
Weiss, L. and Jeltsch, F.: The response of simulated grassland communities
to the cessation of grazing, Ecol. Modell., 303, 1–11,
https://doi.org/10.1016/j.ecolmodel.2015.02.002, 2015.
Wilson, J. B., Peet, R. K., Dengler, J., and Pärtel, M.: Plant species
richness: The world records, J. Veg. Sci., 23, 796–802,
https://doi.org/10.1111/j.1654-1103.2012.01400.x, 2012.
Wurst, S., Sonnemann, I., and Zaller, J. G.: Soil Macro-Invertebrates: Their
Impact on Plants and Associated Aboveground Communities in Temperate Regions
BT – Aboveground–Belowground Community Ecology, in Aboveground-Belowground
Community Ecology, edited by: Ohgushi, T., Wurst, S., and Johnson, S. N., Springer International Publishing,
175–200, 2018.
Zaller, J. G. and Arnone III, J. A.: Earthworm responses to plant species'
loss and elevated CO
2 in calcareous grassland, Plant Soil, 208, 1–8, https://doi.org/10.1023/A:1004424720523, 1999b.
